EUGLENIDS
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CHARACTERISTICS
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They
are single-celled organisms. They carry out all of life´s functions using
only the organelles found in the “typical” eukaryotic cells of animals. Euglenoids
detect light using a pigmented eyespot or stigma red and paraflagellar body
located at the base of the flagellum emerging. All have two flagella
euglenoids rooted basal bodies located in a small reservoir at the front of
the cell. In Euglena, a flagellum is very short and does not protrude from
the cell, while the other is relatively long, and often readily visible under
light microscopy.
Euglena
lacks a cell wall. Instead, has a film formed by a layer of protein on the
support of a substructure of microtubules, spiral strips disposed around the
cell.
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FORM
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They are often
discoidal in shape but can also be ovate, lobate, elongate, U-shaped, or
ribbon-shaped.
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HABITAT
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Most euglenids occur
in fresh water, but a few marine and brackish- water species are known.
Freshwater euglenoids have a contractile vacuole. The majority are noncolonial but some colonial forms exist.
It usually live in quiet ponds or puddles Species of Euglena. They are often abundant in quiet,
inland waters, where they may bloom in numbers sufficient to color the
surface of ponds and ditches green or red.
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SUPPORT AND
LOCOMOTION
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: the shape of
euglenids is maintained by a pellicle consisting of interlocking strips of
protein lying beneath the cell membrane. the stripes that can sometimes be
seen on a euglenid are the seams between the long protein strips winding
around the cell. often the pellicle is suported by regularly arranged
microtubules lying just underneath it. the rigidity of the pellicle is
variable. some have protein strips that are fused together into a rigid
pellicle, while others have protein strips that are articulated to produce a
flexible pellicle. those euglenids with a flexible pellicle undergo euglenoid
movement, or metaboly, in which the cell undulates as it rapidly extends and
contracts . although this type of
movement is not fully understood it is thought to be accomplished by
the sliding of microtubules againsr t the protein strips.
a few euglenids
secrete a lorica, or envelope exterior to the cell membrane . the lorica is
formed by the mucous secretions of small organelles called mucocysts membrane
along the seams between the protein strips of the pellicle. secretions of
mucocysts are also used to form protective coverings when environmental
conditions become unfavorable
locomtion in
euglenids is primarily by flagella. they have two flagella but one may be
very short or represented by just a kinetosome. the flagella originate in an
invagination at the anterior end of the cell called a reservior. the longer,
anteriorly directed flagellum propels the cell through the water or across
surfaces. the shorter flagellum either trails behind for does not emerge from
the reservoir at all. both flagella have a single row of hairs on their
surface and a lattice-like supporting rod, called the paraxonemal rod, lying
adjacent to the microtubules within the shaft.
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NUTRITION
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Euglenids are quite
variable in their nutrition. Aproximately one- third have chloroplasts and
are photo-autotrophic. these are positively phototaxic and have a swelling
near the base of the anterior flagellum that acts as a photoreceptor. the
chloroplasta is surrounded by three membranes and has thylakoids that are
arranged in stacks of three- the photosynthetic pigments include:
chlorophylls a and b, phycobilins, ß-
carotene and the xanthophylls neoxanthin and diadinoanthin.
When feeding as a
heterotroph, the Euglena surrounds a particle of food and consumes it by
phagocytosis.
The cytoplasm of
Euglena and other euglenoids contains many paramylon starch storage granules.
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REPRODUCTION
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Asexual reproduction
in euglenids is by longitudinal cell division. Division occurs along the
longitudinal plane.
Sexual reproduction
has been reported in one species, but this has not been confirmed.
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ECOLOGY
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Euglenids are commonly found in bodies of
water rich in decaying organic matter. As such some of them are useful
indicator organisms of water quality. some species of euglena have been used
in experiments for waste metals such as magnesium, iron and zinc from
sludge. Other euglenids, however, are
environmental pests and some have been shown to produce toxic substances
which have been associated with diseases in trout fry.
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sábado, 22 de junio de 2013
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