sábado, 22 de junio de 2013

EUGLENIDS
CHARACTERISTICS
They are single-celled organisms. They carry out all of life´s functions using only the organelles found in the “typical” eukaryotic cells of animals. Euglenoids detect light using a pigmented eyespot or stigma red and paraflagellar body located at the base of the flagellum emerging. All have two flagella euglenoids rooted basal bodies located in a small reservoir at the front of the cell. In Euglena, a flagellum is very short and does not protrude from the cell, while the other is relatively long, and often readily visible under light microscopy.
Euglena lacks a cell wall. Instead, has a film formed by a layer of protein on the support of a substructure of microtubules, spiral strips disposed around the cell.
FORM
They are often discoidal in shape but can also be ovate, lobate, elongate, U-shaped, or ribbon-shaped.
HABITAT
Most euglenids occur in fresh water, but a few marine and brackish- water species are known. Freshwater euglenoids have a contractile vacuole.  The majority  are  noncolonial but some colonial forms exist. It usually live in quiet ponds or puddles Species of  Euglena. They are often abundant in quiet, inland waters, where they may bloom in numbers sufficient to color the surface of ponds and ditches green or red.
SUPPORT AND LOCOMOTION
: the shape of euglenids is maintained by a pellicle consisting of interlocking strips of protein lying beneath the cell membrane. the stripes that can sometimes be seen on a euglenid are the seams between the long protein strips winding around the cell. often the pellicle is suported by regularly arranged microtubules lying just underneath it. the rigidity of the pellicle is variable. some have protein strips that are fused together into a rigid pellicle, while others have protein strips that are articulated to produce a flexible pellicle. those euglenids with a flexible pellicle undergo euglenoid movement, or metaboly, in which the cell undulates as it rapidly extends and contracts . although this type of  movement is not fully understood it is thought to be accomplished by the sliding of microtubules againsr t the protein strips.
a few euglenids secrete a lorica, or envelope exterior to the cell membrane . the lorica is formed by the mucous secretions of small organelles called mucocysts membrane along the seams between the protein strips of the pellicle. secretions of mucocysts are also used to form protective coverings when environmental conditions become unfavorable
locomtion in euglenids is primarily by flagella. they have two flagella but one may be very short or represented by just a kinetosome. the flagella originate in an invagination at the anterior end of the cell called a reservior. the longer, anteriorly directed flagellum propels the cell through the water or across surfaces. the shorter flagellum either trails behind for does not emerge from the reservoir at all. both flagella have a single row of hairs on their surface and a lattice-like supporting rod, called the paraxonemal rod, lying adjacent to the microtubules within the shaft.
NUTRITION
Euglenids are quite variable in their nutrition. Aproximately one- third have chloroplasts and are photo-autotrophic. these are positively phototaxic and have a swelling near the base of the anterior flagellum that acts as a photoreceptor. the chloroplasta is surrounded by three membranes and has thylakoids that are arranged in stacks of three- the photosynthetic pigments include: chlorophylls a and b, phycobilins,  ß- carotene and the xanthophylls neoxanthin and diadinoanthin.
When feeding as a heterotroph, the Euglena surrounds a particle of food and consumes it by phagocytosis.
The cytoplasm of Euglena and other euglenoids contains many paramylon starch storage granules.
REPRODUCTION
Asexual reproduction in euglenids is by longitudinal cell division. Division occurs along the longitudinal plane.
Sexual reproduction has been reported in one species, but this has not been confirmed.
ECOLOGY
 Euglenids are commonly found in bodies of water rich in decaying organic matter. As such some of them are useful indicator organisms of water quality. some species of euglena have been used in experiments for waste metals such as magnesium, iron and zinc from sludge.  Other euglenids, however, are environmental pests and some have been shown to produce toxic substances which have been associated with diseases in trout fry.

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